Soon after Peter, an orphan, sets sail from England on the ship Never Land, he befriends and assists Molly, a young Starcatcher, whose mission is to guard a trunk of magical stardust from a greedy pirate and the native inhabitants of a remote island.

Fearing that the sinister Lord Ombra was not destroyed, Peter and Molly travel to the land of Rundoon, which is ruled by the evil King Zarboff.

While investigating the strange behavior of the once-peaceful mermaids, the Lost Boys battle their old nemesis, Captain Hook, whose evil plan threatens to destroy the Mollusk village and change life on Neverland Island forever.